Saturday, 13 October 2018

our solar system and life 2

Solar System formation[edit]


An artist's rendering of a protoplanetary disk
The standard model for the formation of the Solar System (including the Earth) is the solar nebula hypothesis.[23] In this model, the Solar System formed from a large, rotating cloud of interstellar dust and gas called the solar nebula. It was composed of hydrogen and helium created shortly after the Big Bang13.8 Ga (billion years ago) and heavier elements ejected by supernovae. About 4.5 Ga, the nebula began a contraction that may have been triggered by the shock wave from a nearby supernova.[24] A shock wave would have also made the nebula rotate. As the cloud began to accelerate, its angular momentumgravity, and inertia flattened it into a protoplanetary disk perpendicular to its axis of rotation. Small perturbations due to collisions and the angular momentum of other large debris created the means by which kilometer-sized protoplanets began to form, orbiting the nebular center.[25]
The center of the nebula, not having much angular momentum, collapsed rapidly, the compression heating it until nuclear fusion of hydrogen into helium began. After more contraction, a T Tauri star ignited and evolved into the Sun. Meanwhile, in the outer part of the nebula gravity caused matter to condense around density perturbations and dust particles, and the rest of the protoplanetary disk began separating into rings. In a process known as runaway accretion, successively larger fragments of dust and debris clumped together to form planets.[25] Earth formed in this manner about 4.54 billion years ago (with an uncertainty of 1%)[26][27][4][28] and was largely completed within 10–20 million years.[29]The solar wind of the newly formed T Tauri star cleared out most of the material in the disk that had not already condensed into larger bodies. The same process is expected to produce accretion disks around virtually all newly forming stars in the universe, some of which yield planets.[30]
The proto-Earth grew by accretion until its interior was hot enough to melt the heavy, siderophile metals. Having higher densities than the silicates, these metals sank. This so-called iron catastrophe resulted in the separation of a primitive mantle and a (metallic) core only 10 million years after the Earth began to form, producing the layered structure of Earth and setting up the formation of Earth's magnetic field.[31] J. A. Jacobs [32] was the first to suggest that the inner core—a solid center distinct from the liquid outer core—is freezing and growing out of the liquid outer core due to the gradual cooling of Earth's interior (about 100 degrees Celsius per billion years[33]).

Hadean and Archean Eons[edit]


Artist's conception of Hadean EonEarth, when it was much hotter and inhospitable to all forms of life.
The first eon in Earth's history, the Hadean, begins with the Earth's formation and is followed by the Archean eon at 3.8 Ga.[2]:145 The oldest rocks found on Earth date to about 4.0 Ga, and the oldest detrital zircon crystals in rocks to about 4.4 Ga,[34][35][36] soon after the formation of the Earth's crust and the Earth itself. The giant impact hypothesis for the Moon's formation states that shortly after formation of an initial crust, the proto-Earth was impacted by a smaller protoplanet, which ejected part of the mantle and crust into space and created the Moon.[37][38][39]
From crater counts on other celestial bodies, it is inferred that a period of intense meteorite impacts, called the Late Heavy Bombardment, began about 4.1 Ga, and concluded around 3.8 Ga, at the end of the Hadean.[40] In addition, volcanism was severe due to the large heat flow and geothermal gradient.[41] Nevertheless, detrital zircon crystals dated to 4.4 Ga show evidence of having undergone contact with liquid water, suggesting that the Earth already had oceans or seas at that time.[34]
By the beginning of the Archean, the Earth had cooled significantly. Present life forms could not have survived at Earth's surface, because the Archean atmosphere lacked oxygen hence had no ozone layer to block ultraviolet light. Nevertheless, it is believed that primordial life began to evolve by the early Archean, with candidate fossils dated to around 3.5 Ga.[42] Some scientists even speculate that life could have begun during the early Hadean, as far back as 4.4 Ga, surviving the possible Late Heavy Bombardment period in hydrothermal vents below the Earth's surface.[43]

Formation of the Moon[edit]


Artist's impression of the enormous collision that probably formed the Moon
Earth's only natural satellite, the Moon, is larger relative to its planet than any other satellite in the solar system.[nb 1] During the Apollo program, rocks from the Moon's surface were brought to Earth. Radiometric dating of these rocks shows that the Moon is 4.53 ± 0.01 billion years old,[46] formed at least 30 million years after the solar system.[47] New evidence suggests the Moon formed even later, 4.48 ± 0.02 Ga, or 70–110 million years after the start of the Solar System.[48]
Theories for the formation of the Moon must explain its late formation as well as the following facts. First, the Moon has a low density (3.3 times that of water, compared to 5.5 for the earth[49]) and a small metallic core. Second, there is virtually no water or other volatiles on the moon. Third, the Earth and Moon have the same oxygen isotopic signature (relative abundance of the oxygen isotopes). Of the theories proposed to account for these phenomena, one is widely accepted: The giant impact hypothesis proposes that the Moon originated after a body the size of Mars (sometimes named Theia[47]) struck the proto-Earth a glancing blow.[1]:256[50][51]
The collision released about 100 million times more energy than the more recent Chicxulub impact that is believed to have caused the extinction of the dinosaurs. It was enough to vaporize some of the Earth's outer layers and melt both bodies.[50][1]:256 A portion of the mantle material was ejected into orbit around the Earth. The giant impact hypothesis predicts that the Moon was depleted of metallic material,[52] explaining its abnormal composition.[53]The ejecta in orbit around the Earth could have condensed into a single body within a couple of weeks. Under the influence of its own gravity, the ejected material became a more spherical body: the Moon.[54]

First continents[edit]

Map with color and texture
Geologic map of North America, color-coded by age. The reds and pinks indicate rock from the Archean.
Mantle convection, the process that drives plate tectonics, is a result of heat flow from the Earth's interior to the Earth's surface.[55]:2 It involves the creation of rigid tectonic plates at mid-oceanic ridges. These plates are destroyed by subductioninto the mantle at subduction zones. During the early Archean (about 3.0 Ga) the mantle was much hotter than today, probably around 1,600 °C (2,910 °F),[56]:82 so convection in the mantle was faster. Although a process similar to present-day plate tectonics did occur, this would have gone faster too. It is likely that during the Hadean and Archean, subduction zones were more common, and therefore tectonic plates were smaller.[1]:258[57]
The initial crust, formed when the Earth's surface first solidified, totally disappeared from a combination of this fast Hadean plate tectonics and the intense impacts of the Late Heavy Bombardment. However, it is thought that it was basaltic in composition, like today's oceanic crust, because little crustal differentiation had yet taken place.[1]:258 The first larger pieces of continental crust, which is a product of differentiation of lighter elements during partial melting in the lower crust, appeared at the end of the Hadean, about 4.0 Ga. What is left of these first small continents are called cratons. These pieces of late Hadean and early Archean crust form the cores around which today's continents grew.[58]
The oldest rocks on Earth are found in the North American craton of Canada. They are tonalites from about 4.0 Ga. They show traces of metamorphism by high temperature, but also sedimentary grains that have been rounded by erosion during transport by water, showing that rivers and seas existed then.[59] Cratons consist primarily of two alternating types of terranes. The first are so-called greenstone belts, consisting of low-grade metamorphosed sedimentary rocks. These "greenstones" are similar to the sediments today found in oceanic trenches, above subduction zones. For this reason, greenstones are sometimes seen as evidence for subduction during the Archean. The second type is a complex of felsic magmatic rocks. These rocks are mostly tonalite, trondhjemiteor granodiorite, types of rock similar in composition to granite (hence such terranes are called TTG-terranes). TTG-complexes are seen as the relicts of the first continental crust, formed by partial melting in basalt.[60]:Chapter 5

Oceans and atmosphere[edit]


Graph showing range of estimated partial pressureof atmospheric oxygen through geologic time [61]
Earth is often described as having had three atmospheres. The first atmosphere, captured from the solar nebula, was composed of light (atmophile) elements from the solar nebula, mostly hydrogen and helium. A combination of the solar wind and Earth's heat would have driven off this atmosphere, as a result of which the atmosphere is now depleted of these elements compared to cosmic abundances.[62] After the impact which created the moon, the molten Earth released volatile gases; and later more gases were released by volcanoes, completing a second atmosphere rich in greenhouse gases but poor in oxygen. [1]:256 Finally, the third atmosphere, rich in oxygen, emerged when bacteria began to produce oxygen about 2.8 Ga.[63]:83–84,116–117
In early models for the formation of the atmosphere and ocean, the second atmosphere was formed by outgassing of volatiles from the Earth's interior. Now it is considered likely that many of the volatiles were delivered during accretion by a process known as impact degassing in which incoming bodies vaporize on impact. The ocean and atmosphere would, therefore, have started to form even as the Earth formed.[64] The new atmosphere probably contained water vapor, carbon dioxide, nitrogen, and smaller amounts of other gases.[65]
Planetesimals at a distance of 1 astronomical unit (AU), the distance of the Earth from the Sun, probably did not contribute any water to the Earth because the solar nebula was too hot for ice to form and the hydration of rocks by water vapor would have taken too long.[64][66] The water must have been supplied by meteorites from the outer asteroid belt and some large planetary embryos from beyond 2.5 AU.[64][67] Comets may also have contributed. Though most comets are today in orbits farther away from the Sun than Neptune, computer simulations show that they were originally far more common in the inner parts of the solar system.[59]:130–132
As the Earth cooled, clouds formed. Rain created the oceans. Recent evidence suggests the oceans may have begun forming as early as 4.4 Ga.[34] By the start of the Archean eon, they already covered much of the Earth. This early formation has been difficult to explain because of a problem known as the faint young Sun paradox. Stars are known to get brighter as they age, and at the time of its formation the Sun would have been emitting only 70% of its current power. Thus, the Sun has become 30% brighter in the last 4.5 billion years.[68] Many models indicate that the Earth would have been covered in ice.[69][64] A likely solution is that there was enough carbon dioxide and methane to produce a greenhouse effect. The carbon dioxide would have been produced by volcanoes and the methane by early microbes. Another greenhouse gas, ammonia, would have been ejected by volcanos but quickly destroyed by ultraviolet radiation.[63]:83

Origin of life[edit]

One of the reasons for interest in the early atmosphere and ocean is that they form the conditions under which life first arose. There are many models, but little consensus, on how life emerged from non-living chemicals; chemical systems created in the laboratory fall well short of the minimum complexity for a living organism.[70][71]
The first step in the emergence of life may have been chemical reactions that produced many of the simpler organiccompounds, including nucleobases and amino acids, that are the building blocks of life. An experiment in 1953 by Stanley Miller and Harold Urey showed that such molecules could form in an atmosphere of water, methane, ammonia and hydrogen with the aid of sparks to mimic the effect of lightning.[72] Although atmospheric composition was probably different from that used by Miller and Urey, later experiments with more realistic compositions also managed to synthesize organic molecules.[73] Computer simulations show that extraterrestrial organic moleculescould have formed in the protoplanetary disk before the formation of the Earth.[74]
Additional complexity could have been reached from at least three possible starting points: self-replication, an organism's ability to produce offspring that are similar to itself; metabolism, its ability to feed and repair itself; and external cell membranes, which allow food to enter and waste products to leave, but exclude unwanted substances.[75]

Replication first: RNA world[edit]

Even the simplest members of the three modern domains of life use DNA to record their "recipes" and a complex array of RNA and protein molecules to "read" these instructions and use them for growth, maintenance, and self-replication.
The discovery that a kind of RNA molecule called a ribozyme can catalyze both its own replication and the construction of proteins led to the hypothesis that earlier life-forms were based entirely on RNA.[76] They could have formed an RNA world in which there were individuals but no species, as mutations and horizontal gene transferswould have meant that the offspring in each generation were quite likely to have different genomes from those that their parents started with.[77] RNA would later have been replaced by DNA, which is more stable and therefore can build longer genomes, expanding the range of capabilities a single organism can have.[78] Ribozymes remain as the main components of ribosomes, the "protein factories" of modern cells.[79]
Although short, self-replicating RNA molecules have been artificially produced in laboratories,[80] doubts have been raised about whether natural non-biological synthesis of RNA is possible.[81][82][83] The earliest ribozymes may have been formed of simpler nucleic acids such as PNATNA or GNA, which would have been replaced later by RNA.[84][85] Other pre-RNA replicators have been posited, including crystals[86]:150 and even quantum systems.[87]
In 2003 it was proposed that porous metal sulfide precipitates would assist RNA synthesis at about 100 °C (212 °F) and at ocean-bottom pressures near hydrothermal vents. In this hypothesis, the proto-cells would be confined in the pores of the metal substrate until the later development of lipid membranes.[88]

Metabolism first: iron–sulfur world[edit]


The replicator in virtually all known life is deoxyribonucleic acid. DNA is far more complex than the original replicator and its replication systems are highly elaborate.
Another long-standing hypothesis is that the first life was composed of protein molecules. Amino acids, the building blocks of proteins, are easily synthesized in plausible prebiotic conditions, as are small peptides (polymers of amino acids) that make good catalysts.[89]:295–297 A series of experiments starting in 1997 showed that amino acids and peptides could form in the presence of carbon monoxide and hydrogen sulfide with iron sulfide and nickel sulfide as catalysts. Most of the steps in their assembly required temperatures of about 100 °C (212 °F) and moderate pressures, although one stage required 250 °C (482 °F) and a pressure equivalent to that found under 7 kilometers (4.3 mi) of rock. Hence, self-sustaining synthesis of proteins could have occurred near hydrothermal vents.[90]
A difficulty with the metabolism-first scenario is finding a way for organisms to evolve. Without the ability to replicate as individuals, aggregates of molecules would have "compositional genomes" (counts of molecular species in the aggregate) as the target of natural selection. However, a recent model shows that such a system is unable to evolve in response to natural selection.[91]

Membranes first: Lipid world[edit]

It has been suggested that double-walled "bubbles" of lipids like those that form the external membranes of cells may have been an essential first step.[92] Experiments that simulated the conditions of the early Earth have reported the formation of lipids, and these can spontaneously form liposomes, double-walled "bubbles", and then reproduce themselves. Although they are not intrinsically information-carriers as nucleic acids are, they would be subject to natural selection for longevity and reproduction. Nucleic acids such as RNA might then have formed more easily within the liposomes than they would have outside.[93]

The clay theory[edit]

Some clays, notably montmorillonite, have properties that make them plausible accelerators for the emergence of an RNA world: they grow by self-replication of their crystalline pattern, are subject to an analog of natural selection (as the clay "species" that grows fastest in a particular environment rapidly becomes dominant), and can catalyze the formation of RNA molecules.[94] Although this idea has not become the scientific consensus, it still has active supporters.[95]:150–158[86]

Cross-section through a liposome
Research in 2003 reported that montmorillonite could also accelerate the conversion of fatty acids into "bubbles", and that the bubbles could encapsulate RNA attached to the clay. Bubbles can then grow by absorbing additional lipids and dividing. The formation of the earliest cells may have been aided by similar processes.[96]
A similar hypothesis presents self-replicating iron-rich clays as the progenitors of nucleotides, lipids and amino acids.[97]

Last universal ancestor[edit]

It is believed that of this multiplicity of protocells, only one line survived. Current phylogenetic evidence suggests that the last universal ancestor (LUA) lived during the early Archean eon, perhaps 3.5 Ga or earlier.[98][99] This LUA cell is the ancestor of all life on Earth today. It was probably a prokaryote, possessing a cell membrane and probably ribosomes, but lacking a nucleus or membrane-bound organelles such as mitochondria or chloroplasts. Like modern cells, it used DNA as its genetic code, RNA for information transfer and protein synthesis, and enzymes to catalyze reactions. Some scientists believe that instead of a single organism being the last universal common ancestor, there were populations of organisms exchanging genes by lateral gene transfer.[98]

Proterozoic Eon[edit]

The Proterozoic eon lasted from 2.5 Ga to 542 Ma (million years) ago.[2]:130 In this time span, cratons grew into continents with modern sizes. The change to an oxygen-rich atmosphere was a crucial development. Life developed from prokaryotes into eukaryotes and multicellular forms. The Proterozoic saw a couple of severe ice ages called snowball Earths. After the last Snowball Earth about 600 Ma, the evolution of life on Earth accelerated. About 580 Ma, the Ediacaran biota formed the prelude for the Cambrian Explosion.[citation needed]

No comments: